LOS ANGELES  — Tim Conway, the impish second banana to Carol Burnett who won four Emmy Awards on her TV variety show, starred aboard “McHale’s Navy” and later voiced the role of Barnacle Boy for “Spongebob Squarepants,” has died. He was 85.

Conway died Tuesday morning in a Los Angeles care facility, according to Howard Bragman, who heads LaBrea Media. Conway’s wife, Charlene Fusco, and a daughter, Jackie, were at his side. The cause was a disorder in which there is an excess of fluid on the brain, Bragman said.

Burnett said in a statement Tuesday that she was heartbroken. “He was one in a million, not only as a brilliant comedian but as a loving human being. I cherish the times we had together both on the screen and off. He’ll be in my heart forever.”

Tributes also came from across the comedy world, including from Conan O’Brien, who said “no one made me laugh harder than Tim Conway” and Kathy Griffin, who called him “a wildly talented, comedy giant.” Al Roker tweeted out a link to Conway playing a hysterically incompetent dentist.

A native of Ohio, Conway credited his Midwestern roots for putting him on the right path to laughs, with his deadpan expression and innocent, simple-minded demeanor.

“I think the Midwest is the heart of comedy in this country, and a little bit of the South, too,” he told the Wisconsin State Journal in 2005. “For some reason, we’re just more laid-back, more understanding. … And Midwesterners have a kinder sense of humor.”

Those qualities probably contributed to his wide popularity on “The Carol Burnett Show,” which he joined in 1975 after years as a frequent guest. The show aired on CBS from 1967 to 1978 and had a short summer stint on ABC in 1979.